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In the void and upon the worlds of Greater Ultramar, the battle for the Imperium continues. Intent on rebuilding his home realm and using it as a base to reconstruct the ravaged stellar empire of mankind, the returned primarch, Roboute Guilliman, proceeds with his war to drive Mortarion and his Death Guard Traitor Legion from the domain of the Ultramarines. But when Guilliman brings his brother to battle upon the diseased plains of Parmenio, the intervention of a greater power in their fraternal struggle threatens to upend the Imperial Regent’s understanding of the galaxy.

As the War for the Realm of Life continues, Lord-Castellant Lorrus Grymn leads the battered remnants of the Steel Souls warrior chamber against the sargasso-citadels of the Verdant Bay. The Hallowed Knights claim victory, but at a terrible cost - Grymn is lost to Realm of Chaos. Now Gardus, newly reforged and fresh from the destruction of the Scabrous Sprawl, must lead his warriors into the foetid heart of Nurgle's realm in search of the Lord-Castellant, where they must once more brave the horrors of the Realm of Decay....